Nikon Z8 Wildlife Photography: Why I Carry Both the NIKKOR Z 400mm f/4.5 VR S and 180-600mm
- 6 days ago
- 10 min read

Following my recent article on the NIKKOR Z 180-600mm, several readers asked the same question: if I also own the NIKKOR Z 400mm f/4.5 VR S, why do I still carry both?
It's a fair question, and one that deserves more than a simple comparison of focal lengths, weight or sharpness.
After countless hours photographing British wildlife with the Nikon Z8, I've come to realise that these lenses aren't competing for a place in my camera bag. Instead, each has earned its place by solving different photographic challenges. The decision isn't about which lens is better, but which one gives me the greatest chance of capturing the images I have in mind before I even leave home.
Every outing begins with that decision. Am I likely to spend the day walking woodland paths in search of small birds, or sitting quietly beside a river waiting for a kingfisher? Will my subjects be predictable, or will they appear unexpectedly at varying distances? These are the questions that influence my choice far more than technical specifications ever could.
This isn't another lens review. There are already plenty of websites that compare charts, measurements and laboratory tests. Instead, I'd like to share how these two exceptional lenses have shaped my own approach to wildlife photography, where each excels, where each falls short, and why, despite the temptation to travel lighter, I've never seriously considered replacing one with the other.
If you're trying to decide between these lenses, or simply wondering why an experienced wildlife photographer would choose to own both, I hope my experiences in the field help answer that question.
Two Lenses, Two Different Mindsets
When photographers ask which lens I prefer, they're often expecting a straightforward answer. The reality is rather different. Long before I consider focal length or maximum aperture, I think about how I want to spend the day.
The NIKKOR Z 400mm f/4.5 VR S encourages me to keep moving. It's a lightweight prime that feels perfectly balanced on the Nikon Z8, making it a pleasure to carry for hours at a time. I find myself walking further, exploring more habitats and reacting instinctively when an opportunity presents itself. Rather than waiting for wildlife to come to me, I'm actively searching for it.
The NIKKOR Z 180-600mm leads me towards a different approach. It offers incredible flexibility, allowing me to adapt instantly as subjects appear at varying distances. Whether I'm watching a kingfisher from across a river, scanning an estuary for wading birds or waiting quietly in a hide, the ability to adjust my composition without moving position is invaluable.
Neither approach is better than the other. They simply suit different situations, and over time I've learnt to recognise which style of photography a location is likely to demand before I even leave home.
That decision has become part of my routine. As I pack my camera bag, I'm not asking myself which lens is the sharper or which reaches further. Instead, I'm thinking about the experience I want from the day ahead. Will I be covering several miles on foot, following woodland trails and responding to fleeting encounters, or will patience be my greatest asset as I wait for wildlife to appear?
Those questions usually provide the answer long before I zip the camera bag closed.

The Lens That Changed the Way I Walk
When Nikon announced the NIKKOR Z 400mm f/4.5 VR S, I'll admit I was intrigued. A lightweight 400mm prime seemed almost too good to be true, particularly for wildlife photography where reach is often everything. What I hadn't appreciated at the time was just how much it would change the way I approached a day in the field.
Before buying the lens, I had become accustomed to carrying heavier telephoto equipment. That's simply part of wildlife photography, and most of us accept it without question. You plan your route more carefully, take more breaks and gradually become aware of the weight on your shoulders as the hours pass.
The first few outings with the 400mm felt noticeably different.
Instead of thinking about conserving energy, I found myself walking a little further.
If a woodland path looked promising, I would follow it. If I heard birds calling deeper into the trees, I was more inclined to investigate. Covering another mile no longer felt like something to avoid, it simply became part of the day's exploration.
That freedom changes your mindset. You're less conscious of the equipment you're carrying and more focused on the environment around you. You begin noticing subtle movements, listening more carefully to birdsong and paying greater attention to changes in the light. The camera becomes less of a burden and more of an extension of the experience.
Of course, a fixed 400mm focal length demands a little more discipline. There are occasions when a bird is simply too close or frustratingly distant, and there's no zoom ring to solve the problem. Instead, you learn to work with your surroundings.
Sometimes that means changing your position, waiting for the subject to move, or accepting that the photograph isn't going to happen that day.
Surprisingly, I've come to enjoy that challenge. It encourages me to think more carefully about composition before raising the camera and to anticipate behaviour rather than reacting to it. Rather than relying on flexibility from the lens, I'm relying on fieldcraft and patience.
Some of my favourite kingfisher, woodland and garden bird photographs have been taken with the 400mm, not because it's technically superior, but because it encourages a slower, more considered way of working. It rewards observation just as much as it rewards good technique.
That's why the 400mm remains my first choice whenever I know I'll be covering ground. It's light enough to carry all day, beautifully balanced on the Nikon Z8 and, perhaps more importantly, it allows me to forget about the equipment and simply enjoy being outdoors.
When Flexibility Becomes More Important Than Weight
As much as I enjoy using the 400mm, there are many occasions when I leave home knowing the NIKKOR Z 180-600mm will give me the best chance of success.
Those decisions are usually made long before I reach my destination, simply by thinking about the type of wildlife I'm hoping to photograph and the environment I'll be working in.
Unlike the 400mm, the 180-600mm is designed to adapt. Wildlife rarely performs to a script, and some of my most memorable encounters have involved birds appearing unexpectedly at completely different distances to those I'd anticipated.
A kingfisher may remain on a favourite perch across the river for several minutes before suddenly diving onto a branch only a few metres away. A heron might take flight without warning and begin circling overhead. Small waders can move constantly as they feed along the shoreline, while birds of prey may appear high above the reserve before disappearing again within seconds.
In these situations, the ability to change focal length instantly becomes invaluable. Rather than worrying about whether the subject will fit comfortably within the frame, I can concentrate entirely on following its behaviour. The zoom becomes almost instinctive, allowing me to respond naturally as the scene unfolds.
That flexibility is particularly valuable when photographing from hides. Space is often limited, your position is fixed and moving closer simply isn't an option.
Whether the birds approach confidently or remain frustratingly distant, the 180-600mm allows me to adapt without disturbing the wildlife or missing the opportunity altogether.
I've also found it to be one of the most forgiving lenses I've ever used. If I arrive somewhere unfamiliar and have little idea how close the birds are likely to be, I know the zoom will cope with almost anything the day presents. That confidence allows me to relax and enjoy the experience rather than worrying whether I've packed the wrong equipment.
Perhaps that's the greatest strength of the 180-600mm. It removes uncertainty. It gives me the freedom to react to whatever nature decides to offer, rather than hoping the conditions match the lens I've chosen.
It's also the lens I recommend most often to photographers building their wildlife kit. Not because it's the sharpest lens Nikon makes, or because it replaces every prime lens, but because it opens the door to an enormous range of photographic opportunities. From small garden birds and kingfishers to deer, dragonflies, birds of prey and distant waders, it's capable of producing exceptional results across an extraordinary variety of subjects.
For me, it remains one of the most versatile wildlife lenses Nikon has ever produced, and despite the arrival of the superb 400mm f/4.5, I wouldn't want to be without it.

Learning to Choose Before I Leave Home
One of the biggest lessons I've learnt over the years is that successful wildlife photography often begins long before I arrive at my destination. Packing the right equipment isn't simply a matter of taking the lens with the greatest reach, it's about understanding the location, the behaviour of the wildlife and, perhaps most importantly, what sort of day I want to have.
If I'm heading to woodland in search of nuthatches, treecreepers or woodpeckers, the 400mm is usually my first choice. I'll be walking for much of the day, stopping frequently to listen, watching for movement among the trees and following whatever opportunities present themselves. The lighter weight makes that style of photography enjoyable, and I rarely finish the day wishing I'd carried something different.
A visit to a nature reserve or an estuary often leads me in the opposite direction. I may spend hours overlooking open water or waiting patiently in a hide where birds can appear at almost any distance. Those are the days when the 180-600mm earns its place. The flexibility to adjust instantly without changing position often means the difference between coming home with a photograph or simply watching the moment unfold.
Weather also plays a part. Strong winds can make smaller birds surprisingly unpredictable, while changing light may encourage me to work differently as the day progresses. During the breeding season I often know where particular species are likely to appear, allowing me to make a more informed decision before leaving home. On other occasions I'm simply exploring somewhere new, and the versatility of the zoom provides reassurance when the unexpected happens.
Experience has taught me that there is rarely a perfect choice. Every lens involves compromise, and accepting those compromises is part of becoming a better wildlife photographer. Rather than trying to carry every possible option, I've learnt to think carefully about the images I'm hoping to create and choose the equipment that best supports that vision.
Of course, there are still days when I wonder if I've packed the wrong lens. That's wildlife photography. Nature has a habit of reminding us that it doesn't follow our plans, and perhaps that's one of the reasons it remains so endlessly rewarding.
Some of my favourite photographs have come from adapting to unexpected situations rather than capturing the images I originally had in mind.
These days I spend less time worrying about whether I've made the perfect equipment choice and more time appreciating the privilege of being outdoors. If I return home having enjoyed the experience, observed something new and perhaps come away with one or two photographs I'm genuinely pleased with, I consider the day a success.
If I Could Only Keep One...
Every photographer eventually asks themselves the same question. If circumstances meant I could only own one of these lenses, which would I keep?
It's not an easy decision because they excel in different ways, and that's precisely why both have remained in my camera bag for so long.
If I wanted a lens that could cope with almost every wildlife encounter, I'd choose the NIKKOR Z 180-600mm. Its versatility is exceptional, and if I were recommending a single lens to someone building a wildlife photography kit around the Nikon Z8, this would almost certainly be my first suggestion. From small garden birds and kingfishers to birds of prey, deer and distant waders, it has repeatedly proven itself to be one of the most adaptable lenses I've ever owned.
But photography isn't always about versatility.
If I knew I would be spending the day walking woodland trails, exploring nature reserves or simply enjoying the freedom to travel light, I'd reach instinctively for the NIKKOR Z 400mm f/4.5 VR S. Every time I pick it up, I'm reminded why it has become such a favourite. It's beautifully balanced on the Nikon Z8, remarkably easy to carry and encourages a style of photography that feels both natural and rewarding.
Perhaps that's the real conclusion.
I don't carry two telephoto lenses because I can't decide which one I prefer. I carry them because each encourages a different way of photographing wildlife. One offers flexibility when nature refuses to follow the script. The other rewards mobility, observation and thoughtful composition. Neither replaces the other because they were never intended to.
After many thousands of photographs, I've stopped asking which lens is better.
Instead, I ask a much simpler question before every outing.
What kind of day do I want to have?
More often than not, the answer to that question tells me exactly which lens to pack.

Further Reading
If you'd like to learn more about my wildlife photography workflow with the Nikon Z8, you may also find these articles and resources useful:
My in-depth review of the NIKKOR Z 180-600mm, where I share extensive field experience using the lens across a wide range of British wildlife subjects.
Photography Gear, including the equipment I use and why I've chosen it.
Nikon Z8 Owner's Guides, with free downloadable resources covering camera setup and field techniques.
Field Journals, where I share the stories behind many of my wildlife photographs, including kingfishers, woodland birds and other British species.
Thank you for taking the time to read this article. I hope it has been useful, whether you're deciding between these two outstanding lenses or simply interested in how I approach wildlife photography in the field. As always, I'd be delighted to hear about your own experiences and which lens you find yourself reaching for most often.
Thank you for reading. If you found this article useful, I'd really appreciate it if you could leave a comment or simply click the like button below. It's always interesting to hear how other photographers approach similar situations, and your feedback helps shape future articles and guides for the website.
